The external structure of mammals is brief. External structure of mammals. Structural features of mammals

Laboratory work No. 10

Issues for discussion

Test yourself

Task 5. Consider the structural features of birds. Indicate the structural features and functions of organ systems and individual organs. Fill out the table. 11 using the textbook “Zoology with elements of ecology” (Blinnikov V.I., pp. 139-146).

Table 11

Characteristics of the structure of birds

What progressive structural features appear in birds compared to reptiles?

Name the adaptations for flight in the internal structure of birds.

Name the structural features of the bird skeleton in connection with adaptation to flight.

Describe the mechanism of double breathing in birds.

What is the structure of a bird egg?

Tasks for independent work

Write down in your notebook the routes of infection and methods of preventing psittacosis. Find out how common psittacosis occurs in the Czech Republic. Use scientific literature and the Internet.

Write down in your notebook three representatives of birds from the Red Book of the Chechen Republic, three representatives from the Red Book of the Russian Federation. Indicate their habitats, reasons for the decline in numbers and ways to restore numbers. Please indicate whether these animals are included in the red list of the International Union for Conservation of Nature (IUCN). For work, use the Internet, electronic versions and originals of the Red Book of the Czech Republic and the Red Book of the Russian Federation.

Target: study the morphological features of mammals

Tasks

Exercise 1. Examine the skeleton of a rabbit. Using fig. 33, find the sections of the spine, determine how the spine of a rabbit, a bird and a lizard differs. Pay attention to the arrangement of the rabbit's limbs compared to the lizard.

Task 2. Consider the shape of mammalian teeth on a wolf's skull. Notice how the teeth are differentiated in shape depending on the function they perform. In Fig. 34, find the main types of teeth.

Task 3. Consider the internal structure of the rat (Fig. 35). Pay attention to the location of the internal organs in the body cavity. Please note regarding large sizes cecum, absence of cloaca and separation anus from genitourinary.

Task 4. Consider the alveoli of the lungs of mammals (Fig. 36). Pay attention to the intensity of entangling of the alveoli with blood vessels.

FEDERAL EDUCATION AGENCY

STATE EDUCATIONAL INSTITUTION

HIGHER STATE EDUCATION

"NOVOSIBIRSK STATE PEDAGOGICAL UNIVERSITY"

FACULTY OF PRIMARY CLASSES

Discipline: Zoology

Structural features and behavioral features of mammals

Performed:

Vashchenko Elena Gennadievna

Novosibirsk 2010

Introduction

Common Mammal Features

Structural features of mammals

Peculiarities of mammalian behavior

Interspecies aggression

Intraspecific aggression

Conclusion

Bibliography

INTRODUCTION

Zoology – scientific discipline that studies animal world, large component biology. Based on the objectives of the study, zoology is divided into a number of disciplines: systematics, morphology, embryology, animal genetics, zoogeography, etc. Based on the objects of research, protozoology, which studies protozoa, is distinguished, invertebrate zoology and vertebrate zoology. The last object of study includes theriology, studying mammals.

The emergence of mammals became possible as a result of the formation of a number of large aromorphoses, which reduced the dependence of animals on changes in the external environment. Mammals evolved from ancient reptiles at the very beginning Mesozoic era, i.e. earlier than birds, but the development that led to the modern richness of forms of this class of vertebrates dates back to the Cenozoic era, after the extinction of large reptiles.

I decided to talk about mammals because... they are the most highly specialized group of land animals. There are currently more than 4,000 species of mammals.

In the first chapter of the essay, I will give an overview of the general features of mammals that distinguish them from other animals, then I will describe the features of their structure and behavior. I will dwell on the behavioral features of mammals in more detail, because... This topic is very interesting and fascinating, but is not covered in the biology textbook.

GENERAL FEATURES OF MAMMALS

Mammals – warm-blooded vertebrates from the amniote group. As I already said, this is the most highly specialized group of land animals, which is distinguished by the following progressive features.

Highly developed central nervous system and sensory organs. The cerebral cortex, formed by gray matter, appears, which ensures a high level of nervous activity and complex adaptive behavior.

Thermoregulation system, ensuring relative constancy of body temperature.

Live birth(except for oviparous ones) and feeding the cubs with mother's milk, which ensures better safety of the offspring.

Height of organization of mammals It is also expressed in the fact that all their organs achieve the greatest differentiation, and the brain has the most perfect structure. The center of higher nervous activity is especially developed in it - the cerebral cortex, consisting of gray brain matter. Due to this reactions and behavior of mammals reach exceptional perfection. This is facilitated by very complex sense organs, especially hearing and smell. The rapid progressive development of mammals was also facilitated by the differentiation of teeth into incisors, canines and molars.

The acquisition played a huge role in the development of this group warm-blooded, that is, a constantly high body temperature. It occurs due to: a) unmixed blood circulation, b) enhanced gas exchange, c) thermoregulatory devices

Unmixed circulation, as in birds, is achieved by a four-chambered heart and the preservation of only one (left) aortic arch in animals. The acquisition of an alveolar lung structure and the appearance of a diaphragm led to increased gas exchange. Diaphragm- This is a muscular septum that completely divides the body into two parts - the thoracic and abdominal. The diaphragm is involved in the act of inhalation and exhalation. Thermoregulation achieved by the appearance of hair and skin glands

Thanks to the perfection of the digestive, respiratory and circulatory systems, the entire metabolism of mammals proceeds very intensively, which, along with high temperature bodies makes them less dependent on climatic conditions environment than amphibians and reptiles. The rapid progressive development of animals is also due to the fact that the highest of them developed viviparity. The embryo is nourished in the womb through a special organ - placenta. After birth, the baby is fed milk. It is secreted by special mammary glands. All this greatly increases the survival rate of offspring.

Thanks to the height of their organization and perfect psyche, by the beginning of the Cenozoic era (65 million years ago), mammals were able to displace the reptiles that had dominated the Earth until then and occupy all the main habitats.

STRUCTURE FEATURES OF MAMMALS

External structure

Animals have well expressed: head, neck, torso and tail. On the head Usually a distinction is made between the cranial region, located behind the eyes, and the facial, or muzzle, located in front. Eyes equipped with upper, lower and third eyelids. Unlike birds, the nictitating membrane (third eyelid) only covers half of the mammal's eye. On the sides of the head there are large ears, at the end of the muzzle there are paired nostrils. Mouth bordered by fleshy lips characteristic of mammals. There are usually very coarse hairs on the upper lip - vibrissae. Several of them are located above the eyes. They play the role of additional organs of touch. Under the root of the tail there is an anal opening, and somewhat anterior to it there is a genitourinary opening. In females, there are 4–5 pairs of nipples on the sides of the body on the ventral side. The limbs are five- or four-fingered, the fingers are armed with claws.

Skin

Wool, covering the body of mammals, is a derivative of the skin. There are two types of hair - guard hair and soft hair - downy hair. The skin consists of two main layers - epidermis And corium. The first is a thin stratum corneum, and the second is very thick and dense. Its lower part forms the subcutaneous tissue.

Skeleton

The spine consists of five sections: cervical, thoracic, lumbar, sacral and caudal. The vertebrae have flat articular surfaces characteristic of mammals and are separated from each other by round cartilaginous discs - menisci.

Cervical region in all mammals (with very rare exceptions) it contains 7 vertebrae. (Both the mouse and the giraffe have 7 cervical vertebrae). These vertebrae lack free ribs. The thoracic region contains 12-13 vertebrae, all of which are equipped with ribs. The front seven pairs of ribs connect to the sternum and are called “true ribs.” The next five pairs do not reach the sternum. The lumbar region is devoid of ribs and usually contains 6-7 vertebrae. The sacrum is formed in most mammals by four fused vertebrae. The anterior ones usually bear two processes, with the help of which the pelvis is articulated. The caudal region is very variable in the number of vertebrae.

Scull is divided into axial, consisting of the bones surrounding the brain, and visceral (facial), which includes the bones surrounding the mouth opening - the palate, the bones of the upper and lower jaws.

Shoulder girdle is represented only by the scapula and clavicle, and mammals do not have a crow bone (coracoid). In fast runners, the collarbone usually disappears (ungulates). The pelvic region consists of a pair of innominate bones, each of which was formed by the fusion of the ilium, ischium and pubis. The skeleton of paired limbs has three typical sections. In the forelimbs these are the shoulder, forearm and hand, and in the hind limbs these are the thigh, lower leg and foot. In mammals, a rounded tendon bone, the patella, appears at the knee joint on the hind limbs.

Muscular system

This system in animals reaches exceptional development and complexity. They have several hundred individual striated muscles. A feature of the mammalian muscular system is the presence of a diaphragm and the appearance of subcutaneous muscles. Diaphragm- This is a dome-shaped muscular septum that separates the thoracic region from the abdominal region. In the center it is perforated by the esophagus. The diaphragm takes part in the acts of breathing and defecation of animals. The subcutaneous muscles represent a continuous subcutaneous layer. With its help, animals can move areas of their skin. The same muscles take part in the formation of lips and cheeks. In monkeys it has almost disappeared and is preserved only on the face. There she received an unusually strong development - this is the so-called facial muscles.

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Nervous system

Brain The beast has powerfully developed hemispheres of the forebrain and cerebellum. They cover all other parts of the brain on top.

Forebrain consists of the cerebral hemispheres covered with gray brain matter - the cerebral cortex. The olfactory lobes extend forward from the hemispheres. Between the hemispheres there is a wide bridge of white nerve fibers.

Diencephalon has a funnel and optic chiasm, as in other classes of vertebrates. The pituitary gland is attached to the funnel of the diencephalon, while the epiphysis is located above the cerebellum on a long stalk. Midbrain It is distinguished by its very small size; in addition to the longitudinal groove, it also has a transverse one, which is characteristic only of mammals. Cerebellum consists of an unpaired part - the vermis and two lateral ones, which are very large and are usually designated as the cerebellar hemispheres. Medulla has a feature that is also characteristic only of mammals. On the sides of this brain are bundles of nerve fibers going to the cerebellum. They are called the posterior cerebellar peduncles. The medulla oblongata passes into the spinal cord.

Sense organs

They are very highly developed in mammals, and, in accordance with the ecological specialization of a particular group, the sense of smell, vision, hearing, and touch are of leading importance. The hearing organs of animals are especially well developed. They have bony ear tympani and large, movable outer ears.

Digestive organs

Oral cavity limited in animals to the lips. The lips are involved in grasping and holding prey. The oral cavity is limited above by a hard bony palate. Due to this, the choanae (inner nostrils) are pushed back towards the pharynx. This allows animals to breathe while food is in the mouth. The sides of the oral cavity are limited by soft muscular cheeks, and at the bottom there is a large muscular tongue. Its functions are to perceive taste sensations and push food under the teeth during chewing and into the pharynx during swallowing. Ducts open into the mouth salivary glands(4 paired glands - parotid, infraorbital, submandibular and sublingual). Teeth do not grow to the surface of the bone, as in previous classes, but sit in independent cells. The teeth are differentiated into incisors, canines and molars. The tooth itself consists of such parts as a crown with a working surface, the body of the tooth and its root. Throat of Beasts short, the windpipe and choanae open into it. Thus, in mammals, the pharynx is the crossroads of two pathways - the alimentary and the respiratory. Esophagus It is a simple, highly extensible muscular tube. After passing through the diaphragm, it connects to the stomach. Stomach has the appearance of a large horseshoe-shaped curved bag that lies across the body. A fat-filled peritoneum hangs from the stomach, which covers all the internal organs like an apron. Liver is located under the diaphragm, its flows open in duodenum, in the loop of which lies the pancreas. Most mammals have gallbladder. Intestines can be of different lengths, it depends on the composition of the feed. The herbivorous rabbit has a very long intestine - 15-16 times longer than the body. Its sections are the small, large and rectal intestines. At the beginning of the large intestine in mammals there is an unpaired blind outgrowth - the cecum. The intestine opens outwards with an independent anal opening.

Respiratory system

Larynx, as usual for mammals, has a cricoid cartilage, in front of which is the large thyroid cartilage. The mammalian larynx is complex. The vocal cords are stretched on the inside of the larynx. These are paired elastic folds of the mucous membrane, stretched in the cavity of the larynx and limiting the glottis. Lungs represent a pair of spongy bodies hanging freely into the chest cavity. Their internal structure is characterized by great complexity. The trachea near the lungs divides into two bronchi. The bronchi, entering the lungs, are divided into secondary bronchi, which in turn are divided into bronchi of the third and fourth order. They end in bronchioles. The ends of the bronchioles are swollen and entwined with blood vessels. These are the so-called alveoli, where gas exchange occurs.

Circulatory system

Heart animals, like birds, have four chambers, and the left ventricle drives blood through the systemic circulation and, like birds, has much thicker walls than the right. A large vessel departs from the left ventricle - the aorta, which begins the systemic circulation. Arterial blood supplies all organs of the body, and venous blood is collected through the vein system. The largest of them - the posterior and two anterior vena cava - flow into the right atrium. From the right atrium, blood enters the right ventricle, from here the pulmonary circulation, or, as it is also called, pulmonary circulation, begins. Venous blood is ejected from the right ventricle into the large pulmonary artery. This artery divides into right and left, leading to the lungs. From each lung, blood collects in the pulmonary vein (the blood in it is arterial), both veins merge and flow into the left atrium. Next, from the left atrium, blood pours into the left ventricle and again flows through the systemic circulation.

Organs, secretions

U mammals are a pair of bean-shaped kidneys located in the lumbar region. From the inner concave side of each kidney there is a ureter (a thin tube) that drains directly into the bladder. The bladder opens into the urethra.

Genitals

In mammals, these are paired testes (in males) or paired ovaries (in females). The testes have a characteristic oval shape. The appendages of the testes are adjacent to them. The paired vas deferens open into the beginning of the urethra. The terminal parts of the vas deferens are expanded into the seminal vesicles. The paired ovaries of the female have an oval-flattened shape. Near each ovary there is an oviduct. At one end the oviduct opens into the body cavity, and at the opposite end it passes into the uterus without a visible border. The uterus of animals is two-horned, the right and left horns of the uterus independently open into the vagina. It is unpaired. The posterior end gradually passes into the urethra and the bladder opens into it. The vagina opens externally through the urogenital opening.

Development of the embryo

Egg cells develop in the ovary, then mature cells exit the ovary into the body cavity and are captured there by the oviduct funnel. Thanks to the flickering movements of the cilia of the tube (oviduct), the egg moves along it, and if the female is fertilized, then in the tube (usually in its first third) the egg and sperm merge. The fertilized egg continues to slowly descend into the uterus and at the same time its fragmentation begins (dividing the egg into many cells). Having reached the uterus, the egg, which by that time has turned into a dense multicellular ball, is embedded in the wall. There, nutrients begin to flow to it. Quite soon, a placenta forms around the implanted embryo. This is the membrane of the fruit, very characteristic of mammals. The placenta is a spongy organ rich in blood vessels, in which a child's and maternal parts are distinguished. The nursery consists of the villi of the embryonic membrane, and the mother's - from the wall of the uterus. During childbirth muscle layer The uterus contracts strongly and the baby's placenta (chorion), by that time connected very slightly with the mucous membrane of the uterus, opens and comes out along with the newborn in the form of a child's place.

BEHAVIORAL FEATURES OF MAMMALS

Intraspecific behavior in mammals is characterized by aggression. It is due to the protection of the species from external and intraspecific factors. Often aggressive behavior manifests itself already in the early stages of ontogenesis, which can lead to the destruction of the youngest cub (kainism), and sometimes to its being eaten by its fellows (cannibalism). Due to aggressive behavior, infanticide (infanticide) is also possible in carnivorous mammals(lions), rodents (gophers), etc. When defending a group territory, collective aggressive behavior of the owners towards strangers is observed. In many cases, aggressive behavior is stimulated by sex hormones. Under the influence of aggressive influence, the body experiences a state of tension, stress. With moderate stress, an increase in autonomic activity is detected nervous system. Stimulation of the adrenal medulla through the autonomic nerves causes them to release adrenaline into the blood. At the same time, changes occur in various parts of the body. The secretion of sweat glands begins, the hair stands on end, the heart beats faster, breathing becomes more frequent and deeper, blood from the digestive tract is redirected to the muscles. All this prepares the body for energetic actions of the required type. Under the influence of chronic stress, the animal becomes ill and may die.

The forms of aggression are diverse, we will try to classify them.

3.1 Interspecies aggression

1. Aggression of the predator towards the prey

In nature, some species inevitably attack others. The mutual influence of predator and prey leads to evolutionary competition, forcing one of them to adapt to the development of the other. But it is worth noting that a predator never completely destroys the prey population; some balance is always established between them. Strictly speaking, ethologists do not consider the behavior of a predator to be aggressive at all (Lorenz, Dolnik, etc.) or consider it a special form of aggression, different from all others. “When a wolf catches a hare, this is not aggression, but hunting. Likewise, when a hunter shoots ducks or a fisherman catches fish, it is not aggressive behavior. After all, they all feel no hostility, no fear, no anger, no hatred towards the victim.” AK. Lorenz writes: “The internal origins of the behavior of a hunter and a fighter are completely different. When a lion kills a buffalo, the buffalo arouses in him no more aggression than does the delicious turkey hanging in the pantry, which I look at with the same pleasure, arouse in me. The difference in internal motivations is clearly visible even in expressive movements. If a dog chases a hare, then it has exactly the same tense and joyful expression with which it greets its owner or anticipates something pleasant. And from the lion’s face, at the dramatic moment of the jump, you can quite clearly see, as is recorded in many excellent photographs, that he is not angry at all. Growling, pinned ears and other expressive movements associated with fighting behavior can be seen in hunting predators only when they are seriously afraid of their armed prey, but even then only in the form of a hint.”

A wonderful illustration of all that has been said is an excerpt from J. London’s story “White Fang”, where a wolf cub, having just dealt with partridge chicks, enters into a fight with the mother partridge. “...He was met by a winged whirlwind. The rapid onslaught and furious blows of the wings blinded and stunned the wolf cub. He buried his head in his paws and squealed. Blows rained down from new strength. The mother partridge was furious. Then the wolf cub got angry. He jumped up with a growl and began to fight back with his paws, then he plunged his small teeth into the bird’s wing and began to pull and drag it from side to side with all his strength. The partridge rushed, hitting him with its other wing. This was the wolf cub's first fight. He was jubilant. He forgot all his fear of the unknown and was no longer afraid of anything. He tore and beat the living creature that struck him. Moreover, this living creature was meat. The wolf cub was possessed by a thirst for blood. He was too absorbed in the fight and too happy to feel his happiness...”

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2. a) Much closer to genuine aggression than the hunter’s attack on the prey, the opposite case is the counterattack of the prey against the predator. An attack on a predator-devourer has obvious meaning for the conservation of the species. Even when the attacker is small, he causes very sensitive troubles to the object of the attack. This is especially true for herd animals, which attack the predator en masse (the so-called mobing). There are many examples. Ungulates often form a tight ring, putting their horns forward and protecting their young. This is how musk oxen repel attacks from wolves, and how buffalos defend themselves from lions. In our domestic cows and pigs, the instinct of a general attack on the wolf is firmly in the blood.

2. b) As when a predator attacks prey or when a predator is baited by its victims, The species-preserving function of the third type of combat behavior, which Lorenz called critical reaction, is also obvious. The expression “fight like a cornered rat” symbolizes a desperate struggle into which the fighter puts everything because he can neither escape nor expect mercy. This form of fighting behavior, the most violent, is motivated by fear (aggression and fear are twins), a strong desire to escape, which cannot be realized because the danger is too close. The animal, one might say, no longer risks turning its back to her - and attacks itself, with the “courage of despair.” This is what happens when escape is impossible due to limited space - as in the case of a cornered rat - but the need to protect the brood or family can do the same. The attack of the mother female on any object that comes too close to the cubs should also be considered a critical reaction. When a dangerous enemy suddenly appears within a certain critical zone, many animals attack him furiously, although they would run from a much greater distance if they noticed his approach from afar.

Besides these special occasions interspecies struggle there are other, less specific ones. Any two animals of different species, approximately equal in strength, can fight over food, shelter, etc. In all of the above cases of struggle between animals there is a common feature: it is quite clear what benefit each of the participants in the battle receives for the preservation of the species. But intraspecific aggression (aggression in the narrow and only sense of the word) also serves to preserve the species, although this is not so obvious.

3.2 Intraspecific aggression

1. Territorial aggression(aggression aimed at protecting territory)

Active protection- an essential sign of territorial behavior. Aggression is manifested towards any member of the same species, especially the same sex. It reaches its maximum at the beginning of the breeding season, when territories are just being established. The territory should not be imagined as a clearly defined space with precisely defined boundaries (it can also be temporary). As a rule, this zone is determined only by the fact that the readiness of a given animal to fight is highest in the place most familiar to it, namely, in the center of its area. That is, the threshold of aggressiveness is lowest where the animal feels most confident, where its aggression is least suppressed by the desire to escape. With distance from this “headquarters”, combat readiness decreases as the situation becomes more alien and fearsome. As one approaches the center of the habitat, aggressiveness increases in geometric progression. This increase is so great that it compensates for all differences in size and strength that may occur in adult sexually mature individuals of the same species.

When the defeated one takes flight, one can observe a phenomenon that occurs in all self-regulating systems with inhibition, namely, oscillations. The pursued, as he approaches his headquarters, regains his courage, and the pursuer, having penetrated into enemy territory, loses his courage. As a result, the fugitive suddenly turns around and - as suddenly as energetically - attacks the recent winner, whom he now beats and drives away. All this is repeated several more times, and, in the end, the fighters stop at a very definite point of balance, where they only threaten each other.

This simple mechanism of struggle for territory ideally solves the problem of a “fair”, that is, the most beneficial for the entire species in its entirety, distribution of individuals across the area in which this type can live. At the same time, even the weaker ones can feed themselves and give birth to offspring, albeit in a more modest space.

Animals can achieve the same effect without aggressive behavior, simply avoiding each other. “Local marking” plays an important role here, especially on the periphery of the site. Feces, secretions of skin glands, optical signs - bark torn from tree trunks, trampled grass, etc. Mammals for the most part “think with their noses,” so it is not surprising that they play a crucial role in marking their possessions with smell. Many mammals leave scent signals in places where they meet or expect to meet a rival. Dogs urinate for this purpose; hyenas, martens, chamois, antelopes and other species use special glands, the secretions of which mark the soil, bushes, stumps, stones, etc. Brown bear scratching his back against a tree, urinating at the same time. Although such marks already hide a silent threat of aggression.

Fight for territory- a very important function of males. Without good land, a family or herd cannot exist; the prosperity of the group depends on their quantity and quality. You need to constantly try to expand your possessions, including at the expense of neighboring groups. Therefore, clashes over territories are inevitable. Human ancestors also lived in territorial groups, and for them the struggle for territory was inevitable. Territory wars for some tribes they became the main occupation in life.

So, taking into account the above, it can be considered reliable that the uniform distribution in space of animals of the same species is the most important function of intraspecific aggression. In addition, Manning highlights another aspect of territorial behavior. Interestingly, in territorial animals, the first reaction of a male to a female contains elements of attack and flight. Such aggressiveness is very important in the formation of “couple attachment.” This aggression, although it can occur between a male and a female, to a large extent redirected outward to neighboring animals. Often a male attacks a male, and a female attacks a female. The cooperation of a male and a female in defending a territory strengthens the bond between them.

2. Marriage tournaments

A certain category of individuals is always involved in mating battles. In most cases, males fight, attacking exclusively or mainly other males of their own species. Sometimes both male and female fight, and when this happens, the male attacks another male, and the female attacks another female. Different species fight differently. First of all, the weapons used are unequal. Dogs bite each other, horses and many other ungulates try to kick their opponents with their forelimbs. Deer measure their strength by locking antlers. What are these collisions for? Charles Darwin already noted that sexual selection - the selection of the best, most powerful animals for procreation - is largely determined by the struggle of rival animals, especially males. The strength of the father provides direct advantages to the offspring in those species where the father takes an active part in caring for the children, especially in their protection. The close connection between the care of males for their offspring and their fights is most clearly manifested in those animals that are not territorial in the above-described sense of the word, but lead a more or less nomadic lifestyle, such as large ungulates, land monkeys, etc. In these animals, intraspecific aggression does not play a significant role in the distribution of space; in the dispersal of species such as bison, various antelopes, horses, which gather in huge communities and to which the division of areas and the struggle for territory are completely alien, because there is plenty of food for them. However, the males of these animals fight each other violently and dramatically, and the selection resulting from this fight results in the emergence of large and well-armed family defenders. This is how such impressive fighters as buffalo bulls or large male baboons arise.

In this regard, it is necessary to mention one more fact - Purely intraspecific selection can lead to the appearance of traits that are not only useless in terms of adaptation to the environment, but also directly harmful to the preservation of the species. Deer antlers, for example, developed exclusively for duels; these antlers are not suitable for anything else. Deer protect themselves from predators only with their front hooves. Such characters develop in cases where selection is directed solely by competition among relatives, without connection with the extraspecific environment. Returning to the topic of the importance of dueling for the preservation of the species, we can say that it serves useful selection only where fighters are tested not only by intraspecific dueling rules, but also by fights with an external enemy. The most important function of a duel is the choice of a fighting defender of the family, thus, another function of intraspecific aggression is to protect the offspring. Proof can be found in the fact that in many animals in which only one sex cares for the offspring, representatives of this particular sex are truly aggressive towards their relatives, or their aggressiveness is incomparably stronger. Something similar is observed in humans.

Aggression in a community of social animals leading to the establishment of hierarchy

Hierarchy- this is the principle of organization, without which, obviously, the ordered joint life of higher animals cannot develop. It consists in the fact that each of the individuals living together knows who is stronger than himself and who is weaker. A relationship of dominance and submission is established in the group, while the number and severity of clashes is reduced because everyone can retreat before the stronger without a fight - and can expect that the weaker, in turn, will retreat in front of him if they get in each other's way. Dolnik emphasizes that victory in skirmishes does not necessarily go to whoever is stronger. It is given to those who are more aggressive: they like to impose conflicts, they threaten a lot and skillfully, and they themselves withstand other people’s threats relatively easily. So, the individual that wins most often becomes dominant. A moment inevitably comes when the dominant takes out his anger on the subdominant (due to a spontaneous outbreak of aggression). He will not answer him, but will redirect the aggression to someone lower on the hierarchical ladder (after all, it’s scary to touch a dominant). By redirecting, aggression will reach those at the lowest level. There is no one to take out aggression on, and it often accumulates. In a large group, there is always a dominant at the top, but there may already be two or three subdominants. This forms a hierarchical pyramid, the bottom layer of which consists of individuals who give in to everyone. They have accumulated a lot of unrealized aggressiveness, hidden by ingratiating behavior in front of their superiors. This is the law of nature and it is impossible to resist it.

Dog-headed monkeys - baboons, hamadryas and others - form hierarchical pyramids based on age. At the head of the group are several older males who have the greatest power and are responsible for the safety of the group (gerontocracy). But, younger males can form alliances and attack superior individuals. Although these alliances are not strong, because the monkeys betray each other all the time, especially when it comes to fighting. Thus, unions can change the hierarchical pyramid through a “revolution from below.” The formation of a pyramid based on age is also characteristic of humans. In traditional societies, the age hierarchy is very strictly observed. But the formation of alliances of subordinates with the aim of overthrowing dominants is also a common thing, known from antiquity to the present day.

The wide distribution of hierarchy convincingly indicates its important species-preserving function: thus, unnecessary struggle between members of the community is avoided. Here the question arises: how is this better than a direct ban on aggression towards members of the community? Ethologists argue that it is impossible to avoid aggression. Firstly, often a community (a wolf pack or a herd of monkeys) urgently needs aggressiveness towards other communities of the same species, so that fighting should be excluded only within the group. Secondly, the tensions that arise within a community as a result of aggressive impulses and the hierarchy that grows out of them can give it much useful structure and strength. The further apart the ranks of two animals are, the less hostility there is between them. And since superior individuals (especially males) necessarily interfere in the conflicts of their inferiors, the principle “The place of the strong is on the side of the weak!” is triggered.

Age hierarchy It also did not arise by chance. With the general progress of evolution, the role of the experience of old animals is increasingly increasing; one might even say that it is joint social life in the most intelligent mammals, due to this, it acquires a new function in the preservation of the species, namely, the traditional transmission of individually acquired information. Naturally, the opposite statement is equally true: joint social life produces selection pressure towards better development of learning abilities, since these abilities in social animals benefit not only the individual, but also the community as a whole. Thus, a long life, significantly exceeding the period of sexual activity, acquires value for the preservation of the species.

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CONCLUSION

In the first chapter of the essay, I examined the main features of the progressive evolution of mammals, which helped them become the dominant animals on the planet. These are three main groups of adaptations: those associated with constant high body temperature; related to the characteristics of reproduction and raising of young; Related big brain in animals of this group. The structure and physiology of animals, and their modern taxonomy are also considered.

In the second chapter, I examined the behavioral features of mammals, in particular interspecific and species aggression. It has been proven that life without aggression is impossible, even if you create an ideal environment that does not contain any irritants. With prolonged failure to perform any instinctive action (manifestation of aggression), the threshold of irritation decreases. A decrease in the threshold of irritation can lead to the fact that, under special conditions, its value can drop to zero, that is, the corresponding instinctive action can “break through” without any external stimulus. In principle, every truly instinctive action, which is deprived of the opportunity to discharge, leads the animal into a state of general anxiety and forces it to search for a discharge stimulus. And a decrease in the irritating threshold and search behavior, rarely in any cases, manifest themselves as clearly as in the case of aggression.

The benefits of aggression can be considered proven. Living space is distributed among animals in such a way that, if possible, everyone finds food for themselves. For the benefit of the offspring, the best fathers and the best mothers are chosen. Children are protected. The community is organized in such a way that a few sophisticated males have sufficient authority so that decisions necessary for the community are not only made, but also carried out. The purpose of aggression is never to destroy a relative, although, of course, during a fight an accident can occur when a horn hits the eye or a fang hits the carotid artery. Aggression is not at all a destructive principle - it is only part of the organization of all living beings, preserving their system of functioning and their very life. Like everything in the world, it can make a mistake - and in doing so destroy a life. However, in the great achievements of the formation of the organic world, this power is intended for good.

BIBLIOGRAPHICAL LIST

1 .Levitin M.G. Biology: Botany. Zoology. Human anatomy and physiology. [Text]/ M.G. Levitin, T.P. Levitin. – St. Petersburg: “Paritet”, 2000;

2 .Lecture in zoology. Features of animals;

3. Dolnik V. Naughty child of the biosphere [Text]/V. Dolnik. - M.: “Pedagogy - press”, 1994;

4.Lorenz K. Aggression: the so-called “evil” [Text]/K. Lorenz. - M.: “Progress”, 1994;

5.Manning O. Animal behavior [Text]/O. Manning. - M.: “Mir”, 1982;

6. Dewsbury, D. Animal behavior: comparative aspects[Text]/D. Dewsbury. - M.: “Mir”, 1981;

7.Fabry, K. Fundamentals of zoopsychology[Text]/K. Fabry. - M.: MSU, 1976;

8. Tinbergen N. Social behavior animals[Text]/N. Tinbergen.- M.: “Mir”, 1993;

9. Bram A. Animal life: Mammals[Text]/A. Brem. - M.: “Terra”, 1992;

10.Tanner ABOUT. Methods of protection in animals [Text]/O. Tenner. - M.: “Mir”, 1985;

11.Biological encyclopedic dictionary [Text]/ M.: “ Soviet encyclopedia", 1989;

12. London J. White Fang [Text]/ J. London. – Ekaterinburg: “Northern House”, 1992.

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FEDERAL EDUCATION AGENCY

STATE EDUCATIONAL INSTITUTION

HIGHER STATE EDUCATION

"NOVOSIBIRSK STATE PEDAGOGICAL UNIVERSITY"

FACULTY OF PRIMARY CLASSES

Discipline: Zoology

Structural features and behavioral features of mammals

Performed:

Vashchenko Elena Gennadievna

Novosibirsk 2010

Introduction

1. General features of mammals

1.2 Structural features of mammals

2. Peculiarities of mammalian behavior

2.2 Interspecies aggression

2.3 Intraspecific aggression

Conclusion

Bibliography

INTRODUCTION

Zoology - a scientific discipline that studies the animal world, a major component of biology. Based on the objectives of the study, zoology is divided into a number of disciplines: systematics, morphology, embryology, animal genetics, zoogeography, etc. Based on the objects of research, protozoology, which studies protozoa, is distinguished, invertebrate zoology and vertebrate zoology. The last object of study includes theriology, studying mammals.

The emergence of mammals became possible as a result of the formation of a number of large aromorphoses, which reduced the dependence of animals on changes in the external environment. Mammals evolved from ancient reptiles at the very beginning of the Mesozoic era, i.e. earlier than birds, but the development that led to the modern richness of forms of this class of vertebrates dates back to the Cenozoic era, after the extinction of large reptiles.

I decided to talk about mammals because... they are the most highly specialized group of land animals. There are currently more than 4,000 species of mammals.

1. GENERAL FEATURES OF MAMMALS

Mammals - warm-blooded vertebrates from the amniote group. As I already said, this is the most highly specialized group of land animals, which is distinguished by the following progressive features.

1. Highly developed central nervous system and sensory organs. The cerebral cortex, formed by gray matter, appears, which ensures a high level of nervous activity and complex adaptive behavior.

2. Thermoregulation system, ensuring relative constancy of body temperature.

3. Live birth(except for oviparous ones) and feeding the cubs with mother's milk, which ensures better safety of the offspring.

Height of organization of mammals It is also expressed in the fact that all their organs achieve the greatest differentiation, and the brain has the most perfect structure. The center of higher nervous activity is especially developed in it - the cerebral cortex, consisting of gray brain matter. Due to this reactions and behavior of mammalsAnddeliver exceptional perfection. This is facilitated by very complex sense organs, especially hearing and smell. The rapid progressive development of mammals was also facilitated by the differentiation of teeth into incisors, canines and molars.

The acquisition played a huge role in the development of this group heatOblood, that is, a constantly high body temperature. It occurs due to: a) unmixed blood circulation, b) enhanced gas exchange, c) thermoregulatory devices

Thanks to the perfection of the digestive, respiratory and circulatory systems, the entire metabolism of mammals proceeds very intensively, which, along with high body temperature, makes them less dependent on environmental climatic conditions than amphibians and reptiles. The rapid progressive development of animals is also due to the fact that the highest of them developed viviparity. The embryo is nourished in the womb through a special organ - placenta. After birth, the baby is fed milk. It is secreted by special mammary glands. All this greatly increases the survival rate of offspring.

2. CCASTRUCTURE FEATURES OF MAMMALS

External structure

Animals have well expressed: head, neck, torso and tail. On the head Usually a distinction is made between the cranial region, located behind the eyes, and the facial, or muzzle, located in front. Eyes equipped with upper, lower and third eyelids. Unlike birds, the nictitating membrane (third eyelid) only covers half of the mammal's eye. On the sides of the head there are large ears, at the end of the muzzle there are paired nostrils. Mouth bordered by fleshy lips characteristic of mammals. There are usually very coarse hairs on the upper lip - vibrissae. Several of them are located above the eyes. They play the role of additional organs of touch. Under the root of the tail there is an anal opening, and somewhat anterior to it there is a genitourinary opening. Females have 4-5 pairs of nipples on the sides of the body on the ventral side. The limbs are five- or four-fingered, the fingers are armed with claws.

Skin

Skeleton

The cervical region of all mammals (with very rare exceptions) contains 7 vertebrae. (Both the mouse and the giraffe have 7 cervical vertebrae). These vertebrae lack free ribs. The thoracic region contains 12-13 vertebrae, all of which are equipped with ribs. The front seven pairs of ribs connect to the sternum and are called “true ribs.” The next five pairs do not reach the sternum. The lumbar region is devoid of ribs and usually contains 6-7 vertebrae. The sacrum is formed in most mammals by four fused vertebrae. The anterior ones usually bear two processes, with the help of which the pelvis is articulated. The caudal region is very variable in the number of vertebrae.

Shoulder girdle is represented only by the scapula and clavicle, and mammals do not have a crow bone (coracoid). In fast runners, the collarbone usually disappears (ungulates). The pelvic region consists of a pair of innominate bones, each of which was formed by the fusion of the ilium, ischium and pubis. The skeleton of paired limbs has three typical sections. In the forelimbs these are the shoulder, forearm and hand, and in the hind limbs these are the thigh, lower leg and foot. In mammals, a rounded tendon bone appears at the knee joint on the hind limbs - the patella.

Muscular system

Nervous system

Brain The beast has powerfully developed hemispheres of the forebrain and cerebellum. They cover all other parts of the brain on top.

Sense organs

Digestive organs

Oral cavity limited in animals to the lips. The lips are involved in grasping and holding prey. The oral cavity is limited above by a hard bony palate. Due to this, the choanae (inner nostrils) are pushed back towards the pharynx. This allows animals to breathe while food is in the mouth. The sides of the oral cavity are limited by soft muscular cheeks, and at the bottom there is a large muscular tongue. Its functions are to perceive taste sensations and push food under the teeth during chewing and into the pharynx during swallowing. The ducts of the salivary glands open into the mouth (4 paired glands - parotid, infraorbital, submandibular and sublingual). Teeth do not grow to the surface of the bone, as in previous classes, but sit in independent cells. The teeth are differentiated into incisors, canines and molars. The tooth itself consists of such parts as a crown with a working surface, the body of the tooth and its root. Throat of Beasts short, the windpipe and choanae open into it. Thus, in mammals, the pharynx is the crossroads of two pathways - the alimentary and the respiratory. Esophagus It is a simple, highly extensible muscular tube. After passing through the diaphragm, it connects to the stomach. Stomach has the appearance of a large horseshoe-shaped curved bag that lies across the body. A fat-filled peritoneum hangs from the stomach, which covers all the internal organs like an apron. Liver located under the diaphragm, its flows open into the duodenum, in the loop of which lies the pancreas. Most mammals have a gallbladder. Intestines can be of different lengths, it depends on the composition of the feed. The herbivorous rabbit has a very long intestine - 15-16 times longer than the body. Its sections are the small, large and rectal intestines. At the beginning of the large intestine in mammals there is an unpaired blind outgrowth - the cecum. The intestine opens outwards with an independent anal opening.

Respiratory system

Circulatory system

Organs, secretions

Genitals

Development of the embryo

Egg cells develop in the ovary, then mature cells exit the ovary into the body cavity and are captured there by the oviduct funnel. Thanks to the flickering movements of the cilia of the tube (oviduct), the egg moves along it, and if the female is fertilized, then in the tube (usually in its first third) the egg and sperm merge. The fertilized egg continues to slowly descend into the uterus and at the same time its fragmentation begins (dividing the egg into many cells). Having reached the uterus, the egg, which by that time has turned into a dense multicellular ball, is embedded in the wall. There, nutrients begin to flow to it. Quite soon, a placenta forms around the implanted embryo. This is the membrane of the fruit, very characteristic of mammals. The placenta is a spongy organ rich in blood vessels, in which a child's and maternal parts are distinguished. The nursery consists of the villi of the embryonic membrane, and the maternal - from the wall of the uterus. During childbirth, the muscular layer of the uterus contracts strongly and the baby's placenta (chorion), by that time connected very slightly with the mucous membrane of the uterus, opens and comes out along with the newborn in the form of a child's place.

3. SPECIALESSENTIALS OF MAMMAL BEHAVIOR

Intraspecific behavior in mammals is characterized by aggression. It is due to the protection of the species from external and intraspecific factors. Often aggressive behavior manifests itself already in the early stages of ontogenesis, which can lead to the destruction of the youngest cub (kainism), and sometimes to its being eaten by its fellows (cannibalism). Due to aggressive behavior, infanticide (infanticide) is also possible in predatory mammals (lions), rodents (gophers), etc. When defending a group territory, collective aggressive behavior of the owners towards strangers is observed. In many cases, aggressive behavior is stimulated by sex hormones. Under the influence of aggressive influence, the body experiences a state of tension, stress. Under moderate stress, increased activity of the autonomic nervous system is detected. Stimulation of the adrenal medulla through the autonomic nerves causes them to release adrenaline into the blood. At the same time, changes occur in various parts of the body. The secretion of sweat glands begins, the hair stands on end, the heart beats faster, breathing becomes more frequent and deeper, blood from the digestive tract is redirected to the muscles. All this prepares the body for energetic actions of the required type. Under the influence of chronic stress, the animal becomes ill and may die.

The forms of aggression are diverse, we will try to classify them.

3.1 Interspecies aggression

1. Aggression of a predator towards its prey

A wonderful illustration of all that has been said is an excerpt from J. London’s story “White Fang”, where a wolf cub, having just dealt with partridge chicks, enters into a fight with the mother partridge. “...He was met by a winged whirlwind. The rapid onslaught and furious blows of the wings blinded and stunned the wolf cub. He buried his head in his paws and squealed. The blows rained down with renewed vigor. The mother partridge was furious. Then the wolf cub got angry. He jumped up with a growl and began to fight back with his paws, then he plunged his small teeth into the bird’s wing and began to pull and drag it from side to side with all his strength. The partridge rushed, hitting him with its other wing. This was the wolf cub's first fight. He was jubilant. He forgot all his fear of the unknown and was no longer afraid of anything. He tore and beat the living creature that struck him. Moreover, this living creature was meat. The wolf cub was possessed by a thirst for blood. He was too absorbed in the fight and too happy to feel his happiness...”

2. b)Like when a predator attacks prey or when baitingschnicknamed by his victims, The species-preserving function of the third type of combat behavior, which Lorenz called critical reaction, is also obvious. The expression “fight like a cornered rat” symbolizes a desperate struggle into which the fighter puts everything because he can neither escape nor expect mercy. This form of fighting behavior, the most violent, is motivated by fear (aggression and fear are twins), a strong desire to escape, which cannot be realized because the danger is too close. The animal, one might say, no longer risks turning its back to her - and attacks itself, with the “courage of despair.” This is what happens when escape is impossible due to limited space - as in the case of a cornered rat - but the need to protect the brood or family can do the same. The attack of the mother female on any object that comes too close to the cubs should also be considered a critical reaction. When a dangerous enemy suddenly appears within a certain critical zone, many animals attack him furiously, although they would run from a much greater distance if they noticed his approach from afar.

In addition to these special cases of interspecies struggle, there are other, less specific ones. Any two animals of different species, approximately equal in strength, can fight over food, shelter, etc. In all of the above cases of struggle between animals there is a common feature: it is quite clear what benefit each of the participants in the battle receives for the preservation of the species. But intraspecific aggression (aggression in the narrow and only sense of the word) also serves to preserve the species, although this is not so obvious.

3.2 Insideid aggression

1. Territorial aggression(aggression aimed at protecting territory)

Active protection- an essential sign of territorial behavior. Aggression is manifested towards any member of the same species, especially the same sex. It reaches its maximum at the beginning of the breeding season, when territories are just being established. The territory should not be imagined as a clearly defined space with precisely defined boundaries (it can also be temporary). As a rule, this zone is determined only by the fact that the readiness of a given animal to fight is highest in the place most familiar to it, namely, in the center of its area. That is, the threshold of aggressiveness is lowest where the animal feels most confident, where its aggression is least suppressed by the desire to escape. With distance from this “headquarters”, combat readiness decreases as the situation becomes more alien and fearsome. As one approaches the center of the habitat, aggressiveness increases exponentially. This increase is so great that it compensates for all differences in size and strength that may occur in adult sexually mature individuals of the same species.

This simple mechanism of struggle for territory ideally solves the problem of a “fair”, that is, the most beneficial for the entire species in its entirety, distribution of individuals across the area in which a given species can live. At the same time, even the weaker ones can feed themselves and give birth to offspring, albeit in a more modest space.

Animals can achieve the same effect without aggressive behavior, simply avoiding each other. “Local marking” plays an important role here, especially on the periphery of the site. Feces, secretions of skin glands, optical signs - bark torn from tree trunks, trampled grass, etc. Mammals for the most part “think with their noses,” so it is not surprising that they play a crucial role in marking their possessions with smell. Many mammals leave scent signals in places where they meet or expect to meet a rival. Dogs urinate for this purpose; hyenas, martens, chamois, antelopes and other species use special glands, the secretions of which mark the soil, bushes, stumps, stones, etc. The brown bear itches its back against a tree, while urinating. Although such marks already hide a silent threat of aggression.

Fight for territory- a very important function of males. Without good land, a family or herd cannot exist; the prosperity of the group depends on their quantity and quality. You need to constantly try to expand your possessions, including at the expense of neighboring groups. Therefore, clashes over territories are inevitable. Human ancestors also lived in territorial groups, and for them the struggle for territory was inevitable. Territorial wars became the main occupation in life for some tribes.

So, taking into account the above, it can be considered reliable that the uniform distribution in space of animals of the same species is the most important function of intraspecific aggression. In addition, Manning highlights another aspect of territorial behavior. Interestingly, in territorial animals, the first reaction of a male to a female contains elements of attack and flight. Such aggressiveness is very important in the formation of “couple attachment.” This aggression, although it can occur between male and female, is largely redirected outward, towards neighboring animals. Often a male attacks a male, and a female attacks a female. The cooperation of a male and a female in defending a territory strengthens the bond between them.

2. Mating tournaments

In this regard, it is necessary to mention one more fact - purely intraspecific selection can lead to the appearance of not only useless traits in terms of adaptation to the environment, but also directly harmful to conservationespecies. Deer antlers, for example, developed exclusively for duels; these antlers are not suitable for anything else. Deer protect themselves from predators only with their front hooves. Such characters develop in cases where selection is directed solely by competition among relatives, without connection with the extraspecific environment. Returning to the topic of the importance of dueling for the preservation of the species, we can say that it serves useful selection only where fighters are tested not only by intraspecific dueling rules, but also by fights with an external enemy. The most important function of a duel is the choice of a fighting defender of the family, thus, another function of intraspecific aggression is to protect the offspring. Proof can be found in the fact that in many animals in which only one sex cares for the offspring, representatives of this particular sex are truly aggressive towards their relatives, or their aggressiveness is incomparably stronger. Something similar is observed in humans.

4. Aregression in the community of social animals, leading to mouthAnew hierarchy

Age hierarchy It also did not arise by chance. With the general progress of evolution, the role of the experience of old animals is increasingly increasing; one can even say that the joint social life of the most intelligent mammals acquires due to this a new function in the preservation of the species, namely, the traditional transmission of individually acquired information. Naturally, the opposite statement is equally true: joint social life produces selection pressure towards better development of learning abilities, since these abilities in social animals benefit not only the individual, but also the community as a whole. Thus, a long life, significantly exceeding the period of sexual activity, acquires value for the preservation of the species.

CONCLUSION

In the first chapter of the essay, I examined the main features of the progressive evolution of mammals, which helped them become the dominant animals on the planet. These are three main groups of adaptations: those associated with constant high body temperature; related to the characteristics of reproduction and raising of young; associated with the large brain in animals of this group. The structure and physiology of animals, and their modern taxonomy are also considered.

BIBLIOGRAPHICAL LIST

1 .Levitin M.G. Biology: Botany. Zoology. Human anatomy and physiology. [Text]/ M.G. Levitin, T.P. Levitin. - St. Petersburg: “Paritet”, 2000;

2 .Lecture in zoology. Features of animals;

3. DolnikIN. Naughty child of the biosphere [Text]/V. Dolnik. - M.: “Pedagogy - press”, 1994;

4. LorenzTO. Aggression: the so-called “evil” [Text]/K. Lorenz. - M.: “Progress”, 1994;

5. ManningABOUT. Animal behavior [Text]/O. Manning. - M.: “Mir”, 1982;

6. Dewsbury, D. Animal behavior: comparative aspects[Text]/D. Dewsbury. - M.: “Mir”, 1981;

7. Fabry, TO. Fundamentals of zoopsychology[Text]/K. Fabry. - M.: MSU, 1976;

8. TinbergenN. Social behavior of animals[Text]/N. Tinbergen.- M.: “Mir”, 1993;

9. BramA. Animal life: Mammals[Text]/A. Brem. - M.: “Terra”, 1992;

10. Tanner ABOUT. Methods of protection in animals [Text]/O. Tenner. - M.: “Mir”, 1985;

11. Biological encyclopedic dictionary [Text]/ M.: “Soviet Encyclopedia”, 1989;

12. LondonJ. White Fang [Text]/ J. London. - Ekaterinburg: “Northern House”, 1992.

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Mammals are the most highly organized vertebrate animals. They are distinguished by a wide variety of sizes and external structure, which depends on conditions and lifestyle. For example, the baby shrew weighs on average 1.5 g, the African elephant weighs 4-5 tons, and the blue whale weighs up to 150 tons.

FEATURES OF THE EXTERIOR STRUCTURE

Let's look at them using a dog as an example. The body of mammals is divided into head, neck, trunk, tail and

two pairs of limbs. The head has an elongated shape. It distinguishes between the cranial and facial sections or muzzle. On the head there is a mouth bordered by movable fleshy lips, above which there is a nose with a pair of nostrils. On the sides of the head there is a pair of eyes protected by movable eyelids. The third eyelid (nictitating membrane) is reduced. Behind the eyes is a pair of movable ears, unique to mammals. The neck provides a movable connection between the head and the elongated body, raised high above the ground on the fore and hind limbs. On its ventral side (in females) there are several pairs of mammary glands, and under the root of the tail there is an anus. The limbs are five-fingered. All fingers end in claws.

COVERS OF THE BODY

Mammalian skin consists of two layers - the epithelial layer and the skin itself. The epithelium is keratinizing. Due to it, sebaceous and sweat glands, hair, claws, nails, horns, and hooves are formed. Presence of hair - characteristic feature mammals. Hair evenly covers the dog's body and is divided into awns (long and coarse), undercoat (short and thin) and downy hairs. The awn protects the skin from damage, and the undercoat serves for thermal insulation. Hair consists of a horny substance - keratin. Due to the seasons, dogs change their coats twice a year.

The sebaceous glands located in the skin produce a secretion that lubricates the surface of the skin and hair, helping to maintain its elasticity, and also protects against the penetration of microorganisms.

Dogs have few sweat glands, because... Their thermoregulation is carried out due to the evaporation of water from the surface of the tongue. The mammary glands are also derivatives of the epidermis, the secretions of which feed the young. In some mammals, sweat or sebaceous glands are modified into odorous ones: musk (muskrat, beaver), anal (predators). Their secret serves for species identification, protection, and marking of the occupied territory.

SKELETON AND MUSCULATE

The skeleton has a structure typical of terrestrial vertebrates, but at the same time a number of features.

The skull is formed by several paired and unpaired fused bones. Its brain section has a larger volume than that of reptiles, which is determined by the significant development of the brain, especially the cortex. The facial section is characterized by the development of a secondary jaw and bony hard palate.

The spinal column consists of 5 sections: cervical, thoracic, lumbar, sacral and caudal. The cervical region consists of 7 vertebrae, which is typical for almost all mammals. The number of thoracic vertebrae ranges from 12 to 15. The ribs are attached to them, which fuse with the sternum to form the rib cage. The massive lumbar vertebrae (6) are movably connected. The sacral vertebrae (3-4) fuse motionlessly between each other and the pelvic bones, creating support for the hind limbs. The caudal region is characterized by great variability in the number of vertebrae.

The belt of the dog's forelimbs is formed by paired shoulder blades and crow bones fused with them. There are no collarbones. The shoulder girdle is connected to the axial skeleton through muscles and ligaments.

The girdle of the hind limbs is formed by paired innominate bones. They are formed as a result of the fusion of the iliac, pubic and ischial bones. Fusing with the sacrum they form a closed pelvis.

The free limbs are five-fingered and have a structure typical of terrestrial vertebrates. The hind limb is characterized by the development of a tendon bone-cup.

The musculature of mammals is highly specialized. The masticatory muscles involved in the capture and grinding of food achieve significant development and differentiation. A characteristic feature of the muscular system is the development of subcutaneous muscles and the diaphragm. The appearance of the diaphragm improves ventilation of the lungs, and also divides the body cavity into the thoracic and abdominal. Subcutaneous muscles play a role not only in thermoregulation, but also in the transmission of information. Good development of the muscles of the limbs ensures greater speed of movement.

INTERNAL STRUCTURE

The digestive system is characterized by the development of specialized teeth, a clear division of the intestinal tube into sections and its considerable length, which ensures effective digestion and absorption of nutrients.

The oral cavity begins with the vestibule of the mouth, the outer wall of which is fleshy lips, and the inner wall is well-developed jaws equipped with specialized teeth.

Dogs have 42 teeth, divided into incisors (12), canines (4), anterior (16) and posterior (10). The teeth have a root, which is strengthened in the jaw socket, and a crown, the shape of which depends

depending on the type of teeth. Dogs' incisors are small, chisel-shaped. The fangs are large, conical, used to capture and kill prey. Molars have wide, tuberous crowns with a sharp cutting edge. The last premolar of the upper jaw and the first molar of the lower jaw form carnassial teeth. In the process of individual development, the milk teeth (incisors, canines and premolars) are replaced by permanent ones.

At the bottom of the oral cavity lies a muscular tongue, the surface of which is covered with taste buds. It is involved in mixing and swallowing food, as well as taste perception. The ducts of three pairs of salivary glands open into the oral cavity, the secretion of which moistens food and also contains enzymes that break down starch.

From the oral cavity, through the pharynx and esophagus, food enters the well-developed simple stomach, and from it, after partial digestion, into the small intestine. The ducts of the liver and pancreas flow into its initial section, the duodenum. Hydrolysis of nutrients and absorption occur in the small intestine. Undigested food remains enter the large intestine, which is divided into the cecum and colon. In these parts of the intestine, feces are formed and removed through the rectum.

RESPIRATORY SYSTEM

Mammals breathe atmospheric air. The main role in gas exchange belongs to the lungs, which are connected to the external environment by the respiratory tract. The respiratory tract includes the nasal cavity, nasopharynx, pharynx, larynx, trachea and bronchi, which form numerous branches in the lungs. The smallest bronchi - bronchioles - end in pulmonary vesicles - alveoli. It is in the latter that gas exchange occurs. In the development of the respiratory organs of mammals, the appearance of the epiglottic cartilage, larynx and alveolar structure of the lung should be noted.

CIRCULATORY SYSTEM

Mammals have a four-chambered heart consisting of two atria and two ventricles. The left aortic arch departs from the left ventricle, unlike in birds. Blood moves through two circles of circulation. The systemic circulation begins from the left ventricle. The arterial blood contained in it is delivered to the tissues through a system of vessels that extend from the aorta. Venous blood collects in the anterior and posterior vena cava, which flow into the right atrium, where the great circle ends.

The pulmonary circulation begins in the right ventricle. From it, venous blood enters the lungs through the pulmonary artery. Oxygenated arterial blood flows from the lungs into the left atrium through four pulmonary veins.

In mammals, due to the development of a four-chambered heart, arterial and venous blood do not mix. Supplying tissues with oxygenated arterial blood enhances redox processes in cells, increasing the level of energy metabolism. As a result, most modern mammals are able to maintain constant temperature body and remain active in conditions of sudden changes in external temperature.

ORGANS OF EXCRETION

In water-salt metabolism in mammals, the main role belongs to the secondary kidneys. They are paired compact bean-shaped bodies located on the sides lumbar region spine. A pair of ureters emerge from the kidneys and open into the bladder, from which urine is discharged through the urethra. The kidneys secrete urine that is hypertonic in relation to blood plasma, which allows saving water by removing metabolic products and salts from the body.

NERVOUS SYSTEM

Consists of the brain, spinal cord and peripheral nerves arising from them. The dog's brain is divided into 5 sections, like all vertebrates, but has a number of features compared to other classes of vertebrates. The forebrain hemispheres reach their greatest size and development. The bulk of them is made up of bark, on the surface of which there is big number convolutions The hemispheres are connected to each other by the corpus callosum.

The midbrain is divided by grooves into a quadrigeminal region, unlike other vertebrates that have a colliculus. Through the anterior colliculus they go to the cortex of the visual tract, and through the posterior colliculus they go to the auditory tract. The cerebellum is large. It consists of hemispheres and a worm located between them. It ensures the maintenance of muscle tone, balance, and coordination of movements. 12 pairs of cranial nerves arise from the brain.

SENSE ORGANS

Well developed. The organs of vision are represented by a pair of eyes. The cornea of the eye is convex, the lens accommodates only due to changes in curvature. In connection with the development of the cortex, secondary associative visual centers are formed in it, located in its occipital lobe.

HEARING ORGAN

It has a complex structure. In the process of evolution, three of its sections were formed: the inner, outer and middle ear. The outer ear is represented by a movable auricle and the external auditory canal. Three auditory ossicles develop in the middle ear: the malleus, the incus, and the stapes. In the inner ear, the cochlea, in which the organ of Corti is located, reaches significant development.

Dogs, like many mammals, have well-developed senses of smell. They are located in the upper-posterior part of the nasal cavity and represent a system of complexly branched shells, the surface of which is covered with olfactory epithelium. The sense of smell allows you to perceive various odors or their combinations characteristic of a group of individuals or an individual.

TASTE ORGANS are represented by taste buds located on the tongue.

SKIN SENSITIVITY is represented by receptors that perceive temperature, pressure, and touch.

GENITAL SYSTEM

Dogs, like all mammals, are dioecious animals. Males have paired testes in which sperm develop. The vas deferens from the testes flow into the urinary canal. The paired ovaries of females lie in the body cavity. One end of the oviduct faces the body cavity, and the other opens into the muscular organ inherent in higher mammals - the uterus, which opens outwards through the vagina.

DEVELOPMENT

Fertilization is internal and occurs in the oviducts. Fertilized eggs, moving along the oviducts, begin to fragment, turning into a multicellular embryo. When the embryo enters the uterine cavity, it attaches to its mucous membrane. At the point of contact of the embryo with the uterine mucosa, the baby's place - the placenta - develops. Through it, during embryonic development, the fetus receives nutrients and oxygen from the mother’s blood, and at the same time removes metabolic products.

Dogs give birth to several blind, helpless cubs. Therefore, parents take care of their offspring. Mothers feed their cubs with milk, keep them warm, and protect them from enemies. And after feeding is completed, the mother and father continue to protect the cubs, raise them, passing on individual experience to their offspring.

MAMMAL BEHAVIOR

The high level of development of the nervous system and sensory organs determines the complexity of mammalian behavior and its plasticity. It is based not only on a set of simple conditioned reflexes, which determine instinctive, innate behavior, but also the ability to form conditioned reflexes and accumulate individual experience on their basis. In the process of interaction of the organism with the environment, there is a constant adaptation of its functional systems to changing conditions based on the formation of new temporary connections in the cerebral cortex and the extinction of old ones. Therefore, the nervous activity of mammals is characterized by mobility, richness and complexity of connections with the environment. Mammals are able to foresee the course of many recurring events and make appropriate decisions in certain situations.

ORIGIN OF MAMMALS

Mammals descend from a group of ancient primitive reptiles - wild-toothed lizards. Based on the remains of the skeletons of beast-toothed lizards, it was established that they lived 200-230 million years ago. Their legs were located under the body and raised it high above the ground. Their teeth had roots and were divided into incisors, canines and molars, and the hard palate was bony, secondary. The skin retained the organizational features of amphibian skin.

Mammals appeared on earth in the Triassic period of the Mesozoic era. Their origin from reptiles is evidenced by common characteristics for both classes: the presence of keratinizing epithelium with homologues of horny scales - hair, the presence of claws on the fingers, homology of the limbs and their girdles, the division of the spinal column into 5 sections, the similarity of the early stages of embryo development. At the same time, mammals have a four-chambered heart and are warm-blooded. They are characterized by feeding their young with milk and viviparity.

The origin of mammals from reptiles is also evidenced by the fact that representatives of the subclass of proto-beasts (platypus, echidna) occupy an intermediate position between reptiles and mammals in structure and reproductive characteristics.

Modern taxonomy divides mammals into 2 subclasses:

1. First beasts and 2. Real beasts. The first subclass includes one order - Monotremes. The second subclass includes the infraclass - Lower animals with the order Marsupials and the infraclass - Higher animals, which unites 19 modern and 12-14 extinct orders.

Both subclasses of mammals originate in the Triassic from the same original group of animal-like reptiles. In subsequent evolution, various adaptations of mammals contributed to their conquest of not only vast areas of land, but also air, soil, fresh and sea waters.

Features of the structure and life processes. Appearance and the sizes of mammals are very diverse depending on conditions and lifestyle. Body weight ranges from 1.5 g (tiny shrew) to 150 tons (blue whale). The long fore and hind limbs are located under the body and facilitate rapid movement, thanks to which animals Not They have no equal in speed of movement. For a cheetah, for example, it reaches 110 km/h.

Leather in mammals it is thicker and more elastic than in animals of other classes. The cells of the outer layer - the epidermis, gradually wearing out and becoming keratinized, are replaced by new, young ones. The inner layer of the skin - the dermis - is well developed, and fat is deposited in its lower part. A derivative of the epidermis are thread-like horny formations - hair. Hair, like the plumage of birds, is a perfect device for thermoregulation. Its basis is made up of fine, soft downy hairs that form the undercoat. Between them, longer, stiffer and sparse guard hairs are developed, protecting downy hair and skin from mechanical damage. In addition, many mammals have long and stiff sensitive hairs - vibrissae - on the head, neck, chest and forelimbs. The hairline changes periodically. The frequency and timing of molting varies among different species of mammals.

Derivatives of the epidermis include nails, claws, hooves, scales and hollow horns (for example, in bulls, goats, rams, antelopes). The bone antlers of deer and elk develop from the inner layer of the skin - the dermis.

have claws (capture prey), fur (adapted to cold conditions), most are small in size due to living in a land-air environment (the most aggressive environment), mimicry, warning coloring, camouflage - protection from enemies, sharp teeth.

54. Internal structure of mammals

Skeleton mammals are basically similar in structure to the skeleton of terrestrial vertebrates, but there are some differences: the number of cervical vertebrae is constant and equal to seven, the skull is more voluminous, which is associated with the larger size of the brain. The bones of the skull fuse quite late, allowing the brain to grow as the animal grows. The limbs of mammals are built according to the five-fingered type, characteristic of terrestrial vertebrates. The methods of movement of mammals are different - walking, running, climbing, flying, digging, swimming - which is reflected in the structure of the limbs. Thus, in the fastest running mammals the number of fingers is reduced: in artiodactyls two (third and fourth) fingers are developed, and in equids - one (third). Animals that lead an underground lifestyle, for example the mole, have an enlarged and uniquely structured hand. Animals capable of gliding (flying squirrels, bats) have elongated phalanges of the fingers and leathery membranes between them.

Digestive system. The teeth sit in the cells of the jaw bones and are divided into incisors, canines and molars. Their number and shape are different and serve as an important systematic feature of animals. In insectivores a large number of poorly differentiated teeth. Rodents are characterized by the strong development of only one pair of incisors, the absence of fangs and the flat chewing surface of the molars. Carnivores have highly developed fangs, which serve to grasp and kill prey, and the molars have cutting chewing tips. Most mammalian species change teeth once in their lifetime. The mouth opening is surrounded by fleshy lips, which is characteristic only of mammals due to milk feeding. In the oral cavity, food, in addition to being chewed by the teeth, is chemically affected by salivary enzymes, and then successively passes into the esophagus and stomach. The stomach in mammals is well separated from other parts of the digestive tract and is equipped with digestive glands. In most mammalian species, the stomach is divided into more or fewer sections. It is most complex in ruminant artiodactyls. The intestine has thin and thick sections. At the border of the thin and thick sections, the cecum emerges, in which fiber is fermented. The ducts of the liver and pancreas open into the cavity of the duodenum. The speed of food digestion is high. Based on the nature of their diet, mammals are divided into herbivores, carnivores and omnivores.

Respiratory system. Mammals breathe light, which have an alveolar structure, due to which the respiratory surface exceeds the body surface by 50 times or more. The breathing mechanism is caused by a change in the volume of the chest due to the movement of the ribs and a special muscle characteristic of mammals - the diaphragm.

Circulatory system There are no fundamental differences between mammals and birds. Unlike birds, in mammals the left aortic arch arises from the left ventricle. In addition, the blood has a high oxygen capacity due to the presence of the respiratory pigment - hemoglobin, contained in numerous small anucleate red blood cells. Due to the high intensity of vital processes and a highly developed thermoregulation system, the body of mammals, like birds, maintains a constant high temperature.

Selection. Mammalian pelvic kidneys are similar By structure with those of birds. Urine with a high content of urea flows from the kidneys through the ureters into the bladder and out of it.

Brain mammals has a relatively large size due to an increase in the volume of the forebrain and cerebellum hemispheres. The development of the forebrain occurs due to the growth of its roof - the cerebral vault, or cerebral cortex.

From sense organs Mammals have better developed senses of smell and hearing. The sense of smell is subtle, allowing you to identify enemies, find food and each other. The organ of hearing in most mammals is quite well developed: in addition to the internal and middle sections, the external auditory canal and the auricle have been formed, which enhances the perception of sounds. In the cavity of the middle ear, in addition to the stapes, like in amphibians, reptiles and birds, in mammals there are two more auditory bones - the malleus and the incus. The sensitive sound-receiving organ of Corti is developed in the inner ear.

Vision for mammals is less significant than for birds. Visual acuity and eye development are different, which is associated with living conditions. Animals that live in open spaces (antelopes) have large eyes and sharp vision, while underground species (moles) have reduced eyes. Function touch perform vibrissae.

Reproduction mammals are characterized by internal fertilization, small eggs (0.05-0.2 mm), devoid of reserve nutrients, viviparity (with the exception of a few species), the construction of special nests by most species for childbirth, as well as feeding newborns with milk.

In most species of mammals, intrauterine development (pregnancy) is associated with the formation of a placenta (or baby's place) in females. Through the placenta, a connection is established between the blood vessels of the child and maternal organisms, which allows for gas exchange in the body of the embryo, the influx of nutrients and the removal of decay products.

The duration of intrauterine development varies among different species: from 11-13 days (in the gray hamster) to 11 months (in the whale). The number of cubs in a litter also varies greatly: from 1 to 12 -15.

A small group of mammals do not develop a placenta and reproduce by laying eggs. But in both cases, the cubs are fed with milk, which contains organic and mineral substances necessary for development.

After breastfeeding is completed, the connection between parents and offspring continues for some time. It is necessary to transmit the individual experience of parents to offspring. Pairs in most mammals are formed for one breeding season, less often for several years (wolves, monkeys).

Origin of mammals. The ancestors of mammals were primitive, unspecialized Paleozoic reptiles - animal-toothed reptiles. Their teeth were differentiated into incisors, canines and molars and were located in cells. In the Triassic, one of the groups of beast-toothed lizards began to acquire the features of a progressive organization and gave rise to mammals.